Akiko Okusu

Photograph of First Last

Senior Lecturer

Contact Info

Phone: 413-545-0480
Office: 140 Morrill II South


B.A., Hamilton College, 1996

Ph.D., Harvard University, 2003


Harvard University, 2004-2005

Research Interests

Invertebrate Evolution and Phylogeny
Evolution Molluscan Bauplan
Molecular Systematics

Representative Publications

Normark, Benjamin B, A. Okusu, G. E. Morse, D. A. Peterson, T. Itioka, and S. A. Schneider. “Phylogeny and Classification of Armored Scale Insects (Hemiptera: Coccomorpha: Diaspididae).” Zootaxa 4616, no. 1 (2019): 1–98.

Schneider, Scott A, A. Okusu, and B. B. Normark. “Molecular Phylogenetics of Aspidiotini Armored Scale Insects (Hemiptera: Diaspididae) Reveals Rampant Paraphyly, Curious Species Radiations, and Multiple Origins of Association with Melissotarsus Ants (Hymenoptera: Formicidae).” Molecular Phylogenetics and Evolution 129 (2018): 291–303.

Peterson, D. A, N. B. Hardy, G. E. Morse, I. C. Stocks, A. Okusu, and B. B. Normark. 2015. Phylogenetic analysis reveals positive correlations between adaptations to diverse hosts in a group of pathogen-like herbivores. Evolution, 69: 2785-2792.

Normark. B. B., G. E. Morse, A. Krewinski, and A. Okusu. 2014. Armored scale insects (Hemiptera: Diaspididae) of San Lorenzo National Park, Panama, with descriptions of two new species. Annals of the Entomological Society of America, 107: 37-49.

Ross, L, N. B. Hardy, A. Okusu, and B. B. Normark. 2013. Large population size predicts the distribution of asexuality in scale insects. Evolution, 67: 196-206.

Sabree, Z. L., C. Y. Huang, A. Okusu, N. A. Moran and B. B. Normark. 2013. The nutrient supplying capabilities of Uzinura, an endosymbiont of armored scale insects. Environmental Microbiology, 15: 1988-1999.

Dunn, C., A. Hejnol, D. Matus, K. Pang, W. Browne, S. Smith, E. Seaver, G. Rouse, M. Obst, G. Edgecombe, M. Sorensen, S. Haddock, A. Schmidt-Rhaesa, A. Okusu, R. Kristensen, W. Wheeler, M. Q. Martindale, and G. Giribet. 2008. Broad taxon sampling improves resolution of the Animal Tree of Life in phylogenomic analyses. Nature, 452: 745-749.

Todt, C., A. Okusu, C. Schander and E. Schwabe. 2008. Aplacophorans and Polyplacophora. In Molluscan Phylogeny by W. Ponder & D.R. Lindberg (eds.). U. C. Berkeley Press, CA, USA. Chapter4. Pp. 71-96.

Giribet, G., A. Okusuz, A.R. Lindgren, S.W. Huff, M. Schrödl and M.K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: Monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA, 103: 7723-7728.

Henry, J. Q., A. Okusu, and M. Q. Martindale. 2004. The cell lineage of the polyplacophoran, Chaetopleura apiculata: variation in the spiralian program and implications for molluscan evolution. Developmental Biology, 272: 145-160.

Okusu, A., E. Schwabe, D. J. Eernisse, and G. Giribet. 2003. Towards a phylogeny of chitons (Mollusca, Polyplacophora) based on combined analysis of five molecular loci. Organisms Diversity and Evolution, 3: 281-302.

Okusu, A. and G. Giribet. 2003. New 18S rRNA sequences from neomenioid aplacophorans and the possible origin of persistent exogenous contamination. J. Moll. Stud., 69: 385-387.

Okusu, A. 2002. Embryogenesis and development of Epimenia babai (Neomeniomorpha Mollusca). Bio. Bull., 203: 87-103.

Reynolds, P. and A. Okusu. 1998. Phylogenetic relationships among families of Scaphopoda (Mollusca). Zoological Journal of the Linnean Society, 126: 131-154.